If that's true then what keeps them from taking over the entire landscape and smothering out all other native understory plants?
I tend to believe that in most situations it's us humans who set the stage for these plants to thrive through our management practices.
In general, it is competition that keeps them from taking over the entire landscape. You are absolutely correct that it is our management techniques in many cases. Here is a great example. I had been planting RR forage soybeans with a light mix of corn each spring to sustain our deer during the summer stress period followed by surface broadcasting a WR/CC/PTT in the fall. It had been working great. Over time, we were able to get our deer numbers in balance enough to use less expensive ag beans.
Then, we did a timber harvest including a thinning and controlled burn of our pines. Marestail was in the native seed bank. It is naturally resistant to glyphosate. It popped up everywhere in the thinned pines. The next spring, I planted soybeans as usual and the fields were inundated by Marestail. Why? It was just as resistant to glyphosate as the soybeans but not attractive to deer like the beans. The marestail was advantaged over beans, there was no other serious competition because of the use of gly in the field, so it actually "canopied" and I ended up with a field of marestail with a few beans.
Over the next few years, natural competition from a variety of other plants as well as pines beginning to close the canopy and dropping needles, eliminated most of the actively growing marestail from the pines. My soybean fields and a couple other new fields that were being created from logging decks where gly was used continued to be inundated with marestail. My clover fields, where incidentally tolerate most "weeds" had a little marestail in it but not enough to be a problem. Fall mowing was enough to keep them in under control.
The soybean fields and new logging decks were another story. I had to stop planting beans. Instead, this year, I sprayed the young marestail with 24D. Because of the soil residual effect, I chose buckwheat for my summer plant. It's large planting window in my area allowed the residual effect of the 24D to dissipate. A thick buckwheat crop is also aggressive enough to smother a lot of competition. I'm making no attempt to control any weeds in this buckwheat. I am hesitant to use gly when I plant the cover crop this fall. Marestail can grow from see or root systems and I don't want to advantage it over other weeds. I would rather have my fall crop compete with a variety of weeds many of which benefit deer and other wildlife much more than marestail.
So, in this example, it was my management techniques that led to the marestail infestation. The use of gly (or any broad spectrum herbicide) in an integral part of most no-till operations. While there are huge long-term benefits to no-till, the use of herbicides can have effects we don't intend and cause their own issues.
Thanks,
Jack